I have spent over 25 years studying varying aspects of DNA replication and genome instability, from in vitro studies of polymerase elongation complexes to the development of methods for whole-genome studies in mammalian systems. In the last 7 years I have been working on developing a pipeline for verifying candidate genes from GWAS studies using mouse models coupled with human and mouse cell model systems.


  • Systems Genetics
  • Biology
  • Biochemistry


  • PhD in Biochemistry, 1992

    University of Virginia

  • BSc in Botany

    University of Iowa